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The raphe is the visible ridge between the lateral halves of the penis, found on the ventral or underside of the penis, running from the meatus opening of the urethra across the scrotum to the perineum area between scrotum and anus.

The human penis differs from those of most other mammals , as it has no baculum or erectile bone and instead relies entirely on engorgement with blood to reach its erect state.

A distal ligament buttresses the glans penis and plays an integral role to the penile fibroskeleton, and the structure is called "os analog," a term coined by Geng Long Hsu in the Encyclopedia of Reproduction.

The human penis cannot be withdrawn into the groin, and it is larger than average in the animal kingdom in proportion to body mass.

Penile measurements vary, with studies that rely on self-measurement reporting a significantly higher average size than those which rely on measurements taken by health professionals.

As of [update] , a systematic review of 15, men and the best research to date on the topic, as the subjects were measured by health professionals concluded that the average length of an erect human penis is Among all primates, the human penis is the largest in girth, but is comparable to the chimpanzee penis and the penises of certain other primates in length.

It was In the developing fetus, the genital tubercle develops into the glans of the penis in males and into the clitoral glans in females; they are homologous.

The urogenital fold develops into the skin around the shaft of the penis and the urethra in males and into the labia minora in females.

On entering puberty , the penis, scrotum and testicles will enlarge toward maturity. During the process, pubic hair grows above and around the penis.

A large-scale study assessing penis size in thousands of to year-old males found no difference in average penis size between year-olds and year-olds.

From this, it can be concluded that penile growth is typically complete not later than age 17, and possibly earlier.

In males, the expulsion of urine from the body is done through the penis. The urethra drains the bladder through the prostate gland where it is joined by the ejaculatory duct , and then onward to the penis.

At the root of the penis the proximal end of the corpus spongiosum lies the external sphincter muscle. This is a small sphincter of striated muscle tissue and is in healthy males under voluntary control.

Relaxing the urethra sphincter allows the urine in the upper urethra to enter the penis properly and thus empty the urinary bladder. Physiologically, urination involves coordination between the central , autonomic , and somatic nervous systems.

In infants, some elderly individuals, and those with neurological injury, urination may occur as an involuntary reflex. Brain centers that regulate urination include the pontine micturition center , periaqueductal gray , and the cerebral cortex.

The distal section of the urethra allows a human male to direct the stream of urine by holding the penis. This flexibility allows the male to choose the posture in which to urinate.

In cultures where more than a minimum of clothing is worn, the penis allows the male to urinate while standing without removing much of the clothing.

It is customary for some men to urinate in seated or crouched positions. The preferred position may be influenced by cultural or religious beliefs. A meta-analysis [24] summarizing the evidence found no superior position for young, healthy males.

For elderly males with LUTS however, in the sitting position compared to the standing:. This urodynamic profile is related to a lower risk of urologic complications, such as cystitis and bladder stones.

An erection is the stiffening and rising of the penis, which occurs during sexual arousal , though it can also happen in non-sexual situations.

Spontaneous erections frequently occur during adolescence due to friction with clothing, a full bladder or large intestine, hormone fluctuations, nervousness, and undressing in a nonsexual situation.

It is also normal for erections to occur during sleep and upon waking. See nocturnal penile tumescence. The primary physiological mechanism that brings about erection is the autonomic dilation of arteries supplying blood to the penis, which allows more blood to fill the three spongy erectile tissue chambers in the penis, causing it to lengthen and stiffen.

The now-engorged erectile tissue presses against and constricts the veins that carry blood away from the penis. More blood enters than leaves the penis until an equilibrium is reached where an equal volume of blood flows into the dilated arteries and out of the constricted veins; a constant erectile size is achieved at this equilibrium.

The scrotum will usually tighten during erection. Erection facilitates sexual intercourse though it is not essential for various other sexual activities.

Although many erect penises point upwards see illustration , it is common and normal for the erect penis to point nearly vertically upwards or nearly vertically downwards or even horizontally straight forward, all depending on the tension of the suspensory ligament that holds it in position.

The following table shows how common various erection angles are for a standing male, out of a sample of 1, males aged 20 through In the table, zero degrees is pointing straight up against the abdomen, 90 degrees is horizontal and pointing straight forward, while degrees would be pointing straight down to the feet.

An upward pointing angle is most common. Ejaculation is the ejecting of semen from the penis, and is usually accompanied by orgasm. A series of muscular contractions delivers semen, containing male gametes known as sperm cells or spermatozoa , from the penis.

It is usually the result of sexual stimulation. Rarely, it is due to prostatic disease. Ejaculation may occur spontaneously during sleep known as a nocturnal emission or wet dream.

Anejaculation is the condition of being unable to ejaculate. Ejaculation has two phases: emission and ejaculation proper.

The emission phase of the ejaculatory reflex is under control of the sympathetic nervous system , while the ejaculatory phase is under control of a spinal reflex at the level of the spinal nerves S2—4 via the pudendal nerve.

A refractory period succeeds the ejaculation, and sexual stimulation precedes it. The human penis has been argued to have several evolutionary adaptations.

The purpose of these adaptations is to maximise reproductive success and minimise sperm competition.

Sperm competition is where the sperm of two males simultaneously resides within the reproductive tract of a female and they compete to fertilise the egg.

This is the process whereby males unwittingly invest their resources into offspring of another male and, evolutionarily speaking, should be avoided.

The most researched human penis adaptations are testis and penis size , ejaculate adjustment and semen displacement.

Evolution has caused sexually selected adaptations to occur in penis and testis size in order to maximise reproductive success and minimise sperm competition.

Sperm competition has caused the human penis to evolve in length and size for sperm retention and displacement. As a result, this adaptation also leaves the sperm less vulnerable to sperm displacement and semen loss.

Another reason for this adaptation is that, due to the nature of the human posture, gravity creates vulnerability for semen loss.

Therefore, a long penis, which places the ejaculate deep in the vaginal tract, could reduce the loss of semen. Another evolutionary theory of penis size is female mate choice and its associations with social judgements in modern-day society.

These varied in height, body shape and flaccid penis size, with these aspects being examples of masculinity.

This is reflected in the association between believed sexual prowess and penis size and the social judgement of penis size in relation to 'manhood'.

Like the penis, sperm competition has caused the human testicles to evolve in size through sexual selection.

The human testicles are moderately sized when compared to other animals such as gorillas and chimpanzees, placing somewhere midway.

Research has also demonstrated that evolutionary adaptations of testis size are dependent on the breeding system in which the species resides.

Human males live largely in monogamous societies like gorillas, and therefore testis size is smaller in comparison to primates in multi-male breeding systems, such as chimpanzees.

The reason for the differentiation in testis size is that in order to succeed reproductively in a multi-male breeding system, males must possess the ability to produce several fully fertilising ejaculations one after another.

One of the primary ways in which a male's ejaculate has evolved to overcome sperm competition is through the speed at which it travels.

Ejaculates can travel up to 30—60 centimetres at a time which, when combined with its placement at the highest point of the vaginal tract, acts to increase a male's chances that an egg will be fertilised by his sperm as opposed to a potential rival male's sperm , thus maximising his paternal certainty.

In addition, males can—and do—adjust their ejaculates in response to sperm competition and according to the likely cost-benefits of mating with a particular female.

The number of sperm in any given ejaculate varies from one ejaculate to another. A male will alter the number of sperm he inseminates into a female according to his perceived level of sperm competition, [29] inseminating a higher number of sperm if he suspects a greater level of competition from other males.

In support of ejaculate adjustment, research has shown that a male typically increases the amount he inseminates sperm into his partner after they have been separated for a period of time.

Increasing the number of sperm a male inseminates into a female acts to get rid of any rival male's sperm that may be stored within the female, as a result of her potential extra-pair copulations EPCs during this separation.

Through increasing the amount he inseminates his partner following separation, a male increases his chances of paternal certainty. This increase in the number of sperm a male produces in response to sperm competition is not observed for masturbatory ejaculates.

Males also adjust their ejaculates in response to sperm competition in terms of quality. Research has demonstrated, for example, that simply viewing a sexually explicit image of a female and two males i.

A female's phenotypic quality is a key determinant of a male's ejaculate investment. Increasing investment in females with high quality phenotypic traits therefore acts to offset the ejaculate investment of others.

Through assessing a female's phenotypic quality, males can judge whether or not to invest or invest more in a particular female, which will influence their subsequent ejaculate adjustment.

The shape of the human penis is thought to have evolved as a result of sperm competition. This means that in the event of a rival male's sperm residing within the reproductive tract of a female, the human penis is able to displace the rival sperm, replacing it with his own.

Semen displacement has two main benefits for a male. Firstly, by displacing a rival male's sperm , the risk of the rival sperm fertilising the egg is reduced, thus minimising the risk of sperm competition.

However, males have to ensure they do not displace their own sperm. It is thought that the relatively quick loss of erection after ejaculation, penile hypersensitivity following ejaculation, and the shallower, slower thrusting of the male after ejaculation, prevents this from occurring.

The coronal ridge is the part of the human penis thought to have evolved to allow for semen displacement.

Research has studied how much semen is displaced by differently shaped artificial genitals. It does this by forcing the semen under the frenulum of the coronal ridge, causing it to collect behind the coronal ridge shaft.

The presence of a coronal ridge alone, however, is not sufficient for effective semen displacement. It must be combined with adequate thrusting to be successful.

It has been shown that the deeper the thrusting, the larger the semen displacement. No semen displacement occurs with shallow thrusting.

The behaviours associated with semen displacement, namely thrusting number of thrusts and depth of thrusts , and duration of sexual intercourse , [50] have been shown to vary according to whether a male perceives the risk of partner infidelity to be high or not.

Males and females report greater semen displacement behaviours following allegations of infidelity. In particular, following allegations of infidelity, males and females report deeper and quicker thrusting during sexual intercourse.

Circumcision has been suggested to affect semen displacement. Circumcision causes the coronal ridge to be more pronounced, and it has been hypothesised that this could enhance semen displacement.

Females report that their vaginal secretions diminish as intercourse with a circumcised male progresses, and that circumcised males thrust more deeply.

The first successful penis allotransplant surgery was done in September in a military hospital in Guangzhou , China.

A recently brain-dead man, aged 23, was selected for the transplant. Despite atrophy of blood vessels and nerves, the arteries , veins , nerves and the corpora spongiosa were successfully matched.

But, on 19 September after two weeks , the surgery was reversed because of a severe psychological problem rejection by the recipient and his wife.

In , researchers Chen, Eberli, Yoo and Atala have produced bioengineered penises and implanted them on rabbits.

This study shows that in the future it could be possible to produce artificial penises for replacement surgeries or phalloplasties. In the world's first successful penis transplant took place in Cape Town , South Africa in a nine-hour operation performed by surgeons from Stellenbosch University and Tygerberg Hospital.

The year-old recipient, who had been sexually active, had lost his penis in a botched circumcision at An Italian nonprofit known as Foregen is working on regrowing the foreskin, with the procedure potentially being partially surgical.

In many cultures, referring to the penis is taboo or vulgar, and a variety of slang words and euphemisms are used to talk about it.

Phallus worship is found in several religions , for example St. Priapus Church [65] and Hindu veneration of the Lingam. The penis is sometimes pierced or decorated by other body art.

Other than circumcision, genital alterations are almost universally elective and usually for the purpose of aesthetics or increased sensitivity.

Piercings of the penis include the Prince Albert , the apadravya , the ampallang , the dydoe , and the frenum piercing.

Foreskin restoration or stretching is a further form of body modification , as well as implants under the shaft of the penis. Trans women who undergo sex reassignment surgery have their penis surgically modified into a vagina via vaginoplasty.

Trans men who undergo such surgery have a phalloplasty. Other practices that alter the penis are also performed, although they are rare in Western societies without a diagnosed medical condition.

Apart from a penectomy , perhaps the most radical of these is subincision , in which the urethra is split along the underside of the penis.

Subincision originated among Australian Aborigines , although it is now done by some in the U. Penis removal is another form of alteration done to the penis.

The most common form of genital alteration is circumcision : removal of part or all of the foreskin for various cultural, religious, and more rarely medical reasons.

For infant circumcision, modern devices such as the Gomco clamp , Plastibell , and Mogen clamp are available. With all modern devices the same basic procedure is followed.

First, the amount of foreskin to be removed is estimated. The foreskin is then opened via the preputial orifice to reveal the glans underneath and ensured that it is normal.

The inner lining of the foreskin preputial epithelium is then separated from its attachment to the glans. The device is then placed this sometimes requires a dorsal slit and remains there until blood flow has stopped.

Finally, part, or all, of the foreskin is then removed. Adult circumcisions are often performed without clamps and require 4 to 6 weeks of abstinence from masturbation or intercourse after the operation to allow the wound to heal.

Advocates of circumcision argue, for example, that it provides important health advantages that outweigh the risks, has no substantial effects on sexual function, has a low complication rate when carried out by an experienced physician, and is best performed during the neonatal period.

The American Medical Association stated in "Virtually all current policy statements from specialty societies and medical organizations do not recommend routine neonatal circumcision, and support the provision of accurate and unbiased information to parents to inform their choice.

From Wikipedia, the free encyclopedia. Human penis A flaccid penis, with surrounding pubic hair removed to show anatomical detail. Main article: Human penis size.

Main article: Development of the reproductive system. See also: List of related male and female reproductive organs.

See also: Tanner stages. Main article: Urination. Main article: Erection. Main article: Ejaculation.

Main article: Genital modification and mutilation. Main article: Circumcision. Dissection showing the fascia of the penis as well as several surrounding structures.

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